When users want to access the site, they will send an http request to the hosted server, and the server will respond by displaying the website's content. This process will follow protocols and network architectures. User's will be able to connect to the site from all around the world, due to the global connectivity of the internet.
The user's that use the desktop application will also tap into the server, but not using the http protocol. The application will most likely request information from the information database, which will be located on the server. In this sense, the information is the same as the web version, but it's acquiring it in a different way.
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